AB, Bowdoin College; MEM, Yale University; PhD, University of Wyoming
Wildlife ecology, management, and policy; Biodiversity conservation; Science outreach and communication
My group studies the effects of environmental change on the behavioral, population, and community ecology of wide-ranging wildlife. We are especially interested in the development of solutions to human-wildlife conflicts. We work with agency, university, non-profit, and private partners to make sure we are asking the right questions and making the best research contributions we can. We also work with photographers, filmmakers, artists, and others to communicate about science and conservation to the public. Our work is motivated by a global need for science and conservation solutions that promote both ecological integrity and human wellbeing, especially in working landscapes. Read more about our work on the Middleton Lab website.
Middleton, A.D., T. Stoellinger, H. Karandikar, B. Leonard, H. Doremus, and C. Kremen. In review. Harnessing visitors' enthusiasm for national parks to advance transboundary conservation.
Smith J.A., E. Donadio, O.R. Bidder, J.N. Pauli, M.J. Sheriff, and A.D. Middleton. In revision. Temporal flexibility enables an ambush carnivore to balance prey encounter rate and capture ability.
Middleton, A.D., H. Sawyer, J.A. Merkle, M.J. Kauffman, E.K. Cole, S.R. Dewey, J.A. Gude, D.D. Gustine, D.E. McWhirter, K.M. Proffitt, and P.J. White. Accepted. Conserving ungulate migrations requires transboundary science, policy, and management. Frontiers in Ecology and the Environment.
Sawyer, H., C. W. Lebeau, T. L. MacDonald, W. Xu, and A.D. Middleton. 2019. All routes are not created equal: An ungulate’s choice of migration route can influence its survival. Journal of Applied Ecology 56:1860-69.
Smith, J.A., E. Donadio, J.N. Pauli, M.J. Sheriff, and A.D. Middleton. 2019. Habitat complexity mediates the predator-prey space race. Ecology 100:e02724.
Rickbeil G.J.M, J.A. Merkle, G. Anderson, M.P. Atwood, J.P. Beckmann, E.K. Cole, A.B. Courtemanch, S. Dewey, D.D. Gustine, M.J. Kauffman, D.E. McWhirter, T. Mong, K. Proffitt, D.R. Stahler, P.J. White, and A.D. Middleton. 2019. Plasticity in elk migration timing is a response to changing environmental conditions. Global Change Biology 25:2368-2381.
Smith, J.A., E. Donadio, J.N. Pauli, M.J. Sheriff, and A.D. Middleton. 2019. Integrating temporal refugia into landscapes of fear: prey exploit predator downtimes to forage in risky places. Oecologia 189:883-890.
Gaynor, K.M. J.S. Brown, A.D. Middleton, M.E. Power, and J.S. Brashares. 2019. Landscapes of fear: spatial patterns of risk perception and response. Trends in Ecology and Evolution 34: 355-368.
Sawyer, H., J.A. Merkle, A.D. Middleton, S.P.H. Dwinnell, and K. Monteith. 2019. Migratory plasticity is not ubiquitous in large herbivores. Journal of Animal Ecology 88:450-460.
van Eeden, L. M., A. Eklund, J. R. B. Miller, J. V. López-Bao, G. Chapron, M. R. Cejtin, M. S. Crowther, C. R. Dickman, J. Frank, M. Krofel, D. W. Macdonald, J. McManus, T. K. Meyer, A. D. Middleton, T. M. Newsome, W. J. Ripple, E. G. Ritchie, O. J. Schmitz, K. J. Stoner, M. Tourani, A. Treves. 2018. Carnivore conservation needs evidence-based livestock protection. PLoS Biology 16:e2005577.
Middleton, A.D., J.A. Merkle, D.E. McWhirter, J.G. Cook, R.C. Cook, P.J. White, and M.J. Kauffman. 2018. Green-wave surfing increases fat gain in a migratory ungulate. Oikos 127:1060-68.
Lukacs, P., M. Hebblewhite, B. Johnson, H. Johnson, M. Kauffman, M. Mitchell, K. Proffitt, P. Zager, J. Brodie, K. Hersey, A. Holland, M. Hurley, S. McCorquodale, A.D. Middleton, M. Nordhagen, J. Nowak, D. Walsh, P.J. White. 2018. Factors influencing elk recruitment across three ecotypes in the western USA. Journal of Wildlife Management 82:698-710.
Perrig, P., E. Donadio, A.D. Middleton, and J.N. Pauli. 2017. Predation subsidizes an obligate scavenger in the high Andes. Journal of Applied Ecology.
Sawyer, H., A.D. Middleton, K. Monteith, M. Hayes, and M.J. Kauffman. 2016. The extra mile: Ungulate migration distance alters use of seasonal range and exposure to anthropogenic risk. Ecosphere 7:e01534.
Merkle, J.A., K.L. Monteith, E.O. Aikens, M.M. Hayes, K.R. Hersey, A.D. Middleton, B.A. Oates, H. Sawyer, B.M. Scurlock, and M.J. Kauffman. 2016. Large herbivores surf waves of green-up in spring. Proceedings of the Royal Society of London B 283:20160456.
Nelson, A.A., M.J. Kauffman, A.D. Middleton, M.D. Jimenez, D.E. McWhirter, and K. Gerow. 2015. Native prey distribution and migration mediates wolf predation on domestic livestock in the Greater Yellowstone Ecosystem. Canadian Journal of Zoology 94:291-299.
Middleton, A.D., M.J. Kauffman, D.E. McWhirter, R.C. Cook, J.G. Cook, M.D. Jimenez, S.E. Albeke, H. Sawyer, and P.J. White. 2013. Linking antipredator behavior to prey demography reveals the limited risk effects of an actively hunting large carnivore. Ecology Letters 16:1023-1030.
Middleton, A.D., T.A. Morrison, J.K. Fortin, M.J. Kauffman, C.T. Robbins, K.M. Proffitt, P.J. White, D.E. McWhirter, T.M. Koel, D. Brimeyer, and W.S. Fairbanks. 2013. Grizzly bear predation links the loss of native trout to the demography of migratory elk in Yellowstone. Proceedings of the Royal Society B – Biological Sciences 280:20130870.
Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. 2013. Rejoinder: challenge and opportunity in the study of ungulate migration amid environmental change. Ecology 94:1280-86.
Middleton, A.D., M.J. Kauffman, D.E. McWhirter, J.G. Cook, R.C. Cook, A.A. Nelson, M.D. Jimenez, and R.W. Klaver. 2013. Animal migration amid shifting patterns of predation and phenology: Lessons from a Yellowstone elk herd. Ecology 94:1245-56.
Sawyer, H. M.J. Kauffman, A.D. Middleton, T.A. Morrison, R.M. Nielson, and T.B. Wyckoff. 2013. A framework for understanding barrier effects on migratory ungulates. Journal of Applied Ecology 50:68-78.
Nelson, A.A., M.J. Kauffman, A.D. Middleton, M.D. Jimenez, D.E. McWhirter, J.J. Barber, and K. Gerow. 2012. Elk migration patterns and human activity influence wolf habitat selection in the Greater Yellowstone Ecosystem. Ecological Applications 22:2293-2307.
Cross, P.C., E.K. Cole, A.P. Dobson, W.H. Edwards, K.L. Hamlin, G. Luikart, A.D. Middleton, B.M. Scurlock, and P.J. White. 2010. Probable causes of increasing elk brucellosis in the Greater Yellowstone Ecosystem. Ecological Applications 20:278-288.
POPULAR (NON-ACADEMIC) WRITING
Middleton, A.D. 2019. Tracking Wolves, pp. 179-187 in "Matthew Barney, Redoubt" by Yale University Art Gallery, Yale University Press, New Haven, CT. 368 pp.
Middleton, A.D. 2017. Sustaining migrations in the modern West. In “Yellowstone Migrations" by J. Riis. Pp 119-124.
Middleton, A.D. and L. Allison. 2016. Beyond boundaries in the Greater Yellowstone Ecosystem: scientists and ranchers exploring solutions for the future. Western Landowners Alliance. 28pp.
Middleton, A.D. 2016. Seeing Yellowstone: Science at its limits in a conservation crucible. pp. 36-51 in “Invisible Boundaries: Exploring Yellowstone’s Great Animal Migrations” ed. Karen B. McWhorter. Buffalo Bill Center of the West, Cody, WY. 80pp.
Middleton, A.D., C.R. Preston and K. McWhorter, co-curators. 2016. "Invisible Boundaries: Exploring Yellowstone's Great Animal Migrations." Buffalo Bill Center of the West and National Geographic Society.
I currently serve as a Fellow of the National Geographic Society and a Trustee of the Buffalo Bill Center of the West.
130 Mulford Hall #3114
Berkeley, CA 94720